Sarina Seyedein - Deptartment of Plant Protection, Faculty of Plant Production, Gorgan University of Agricultural Science and Natural Resource, Golestan, Iran.
Vahid Rahiminejad - Deptartment of Plant Protection, Faculty of Plant Production, Gorgan University of Agricultural Science and Natural Resource, Golestan, Iran.
Ahmad Nadimi - Deptartment of Plant Protection, Faculty of Plant Production, Gorgan University of Agricultural Science and Natural Resource, Golestan, Iran.

During a comprehensive survey of the heterostigmatic mites associated with dung beetles in the hyrcanian forests, northern Iran, several colonies of the pygmephoroid mites including four species of the genus Spatulaphorus Rack, ۱۹۹۳, S. copridis Khaustov, ۲۰۰۷, S. gorganica Rahiminejad & Hajiqanbar, ۲۰۱۱, S. vladimiri Khaustov, ۲۰۰۵ and S. geotruporum Khaustov, ۲۰۰۵ phoretic on Geotrupes spiniger (Marsham, ۱۸۰۲) were found. This is the first record of S. geotruporum from Asia. Furthermore, this is the first record of phoresy of S. geotruporum on G. spiniger.

Heterostigmatina, Dung beetles, Phoresy, Host preference, Iran, هترواستیگماتینا, سرگین غلطان, همسفری, ترجیح میزبانی, ایران